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Plot
Season one
Main article: Teenage Mutant Ninja Turtles (2012 TV series) (season 1)
Ninjutsu master Hamato Yoshi (Splinter) is carrying his four pet turtles through the streets of Manhattan when he encounters members of an alien race called the Kraang. During an altercation with these aliens, Yoshi and the turtles are exposed to the Kraang's chemical, called mutagen, which causes organic beings to undergo major physical transformations. Yoshi takes on characteristics of a rat, and the turtles take on human characteristics. Yoshi retreats to the New York City sewers where he raises the four turtles as his sons and imparts to them his knowledge of ninjutsu.
Now teenagers, the turtles (Leonardo, Raphael, Donatello and Michelangelo) venture to the surface for the first time and learn that the Kraang are using the mutagen as part of their plan to take over New York City. They befriend the teenage April O'Neil after she and her psychologist father Kirby are abducted by the Kraang. Donatello, who has developed a crush on April, is able to rescue her and while Kirby remains a prisoner of the Kraang, she becomes an ally of the turtles, who try to help her locate her father. April is also trained by Splinter to be a kunoichi, a female ninja.
After learning of Splinter's and the turtles's presence in New York, Splinter's adoptive brother and long-time enemy Oroku Saki (Shredder) travels there and orders his birth clan, the Foot to track down Splinter and his sons, and put an end to their clan. This family feud eventually earns the coincidence of two of Shredder's disciples, martial arts star Chris Bradford and Brazilian street thug Xever Montes, getting mutated into Dogpound (later Rahzar) and Fishface. Learning about the Kraang's presence through his adopted daughter Karai, Shredder enters an alliance with the aliens to destroy their mutual enemies in the Hamato Clan.
The turtles later discover that the Kraang came to Earth from Dimension X and built their headquarters with a company called T.C.R.I and are plotting to use the mutagen and a set of Kraang-like powers that April was born with to convert Earth into a planet suitable for their race. After the turtles rescue Kirby, the Kraang invade New York, but the turtles and April emerge triumphant when they send the Technodrome, the Kraang leader Kraang Prime's ship, crashing into the sea. Meanwhile, Splinter battles Shredder and learns that Karai is his own daughter, Hamato Miwa, who was kidnapped by Shredder and who believes that Splinter killed her mother. The season ends with turtles celebrating their victory over the Kraang as Splinter hides the secret of Miwa from them.
Season two
Main article: Teenage Mutant Ninja Turtles (2012 TV series) (season 2)
The turtles struggle to contain an outbreak of mutations that occurs thanks to the leftover mutagen from the thwarted Kraang invasion. Kirby is among the victims of the outbreak, and a misunderstanding leads April to became very upset and ashamed of the turtles and break off their friendship. However, the turtles are able to earn her forgiveness when they save her from Karai, who has taken temporary command of the Foot while the Shredder is away in Japan. Along with April came her new friend Casey Jones, who helps repel an assault on the turtles' lair. Kirby is eventually restored to his human form when Donatello manages to concoct another Kraang chemical called retro-mutagen, which causes organic beings to reverse their major physical transformations, and help keep it away from former T.C.R.I inventor Baxter Stockman, who gets mutated into Stockman-Fly while under Shredder's employ.
Meanwhile, Shredder returns from Japan with the mutated Japanese bounty hunter Tiger Claw as his new second-in-command. During a battle with the turtles, Karai is informed of her true nature by Leonardo (who has developed a crush on her since season 1), but she is too reluctant to accept it. When she is taken to the lair, she finally realizes the truth and disowns Shredder, who responds by imprisoning her. Repeated attempts to free Karai ultimately succeed, but Shredder captures her again and uses her as bait to kill the Hamato Clan. However, Karai ends up mutated into a serpent, to everyone's horror.
In response to this, Shredder has the Foot Clan help the Kraang, who have just perfected the previously unstable mutagen, launch a second invasion of New York, starting by destroying the turtles' lair and forcing them to abandon it. The Kraang begin mutating New York's populace, including Kirby, despite the efforts of Earth's military and the turtles. Leonardo is gravely wounded when he is ambushed by the entire Foot, and Splinter is seemingly killed during a battle with Shredder. The turtles, April, and Casey are forced to flee New York as the Kraang successfully conquer the city.
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